What is the result of applying the signed reversal ρ (B, D) in the circular genome π = (ABCD) (D -B -C -A)?

1. ρπ = (B C -A -D) (D A -C -B)
2. ρπ = (A B C D) (-D -C -B -A)
3. ρπ = (B C -D -A) (-C -B A D)
4. ρπ = (-D -C -B A) (-A B C D)
5. None of the above.

Original idea by: Marcus Vinícius Benedito
Translate help: Leandro José de bortoli